400mm =Super-Telephoto Prime
f/4.5= Constant Aperture
1,160 g¹ =Lightweight
23.45 cm= Compact Size
5.5 stops⁴ =Powerful Vibration Reduction  (VR)
Light. Compact. S-Line Performance.
Take in the distance with this compact and agile super-telephoto prime lens. The NIKKOR Z 400mm f/4.5 VR S provides the reach that you need, and yet is nimble enough for handheld shooting. Acquire clear and crisp images with its powerful Vibration Reduction (VR) and superior S-Line optical performance as you photograph the action in full speed from afar.
Handheld shooting made easy.
Photograph distant wildlife and subjects while staying light on the move. Weighing at approximately 1,160g¹, the NIKKOR Z 400mm f/4.5 VR S is the lightest² lens in its class. With its size at 234.5mm (L) x 104mm (D), shooting handheld is even easier. 
So far, yet so close.
With a super-telephoto focal length of 400mm, you can capture every delicate detail of a dolphin breaking the surface, or a squirrel devouring an acorn. Take it even further by attaching either the Z TELECONVERTER TC-1.4x or Z TELECONVERTER TC-2.0x³ to extend the focal length to 560mm or 800mm respectively, so you can get close to your subject without disturbance.
Powerful Vibration Reduction (VR)
Stay sharp with the NIKKOR Z 400mm f/4.5 VR S. Its in-lens Vibration Reduction (VR) compensates up to 5.5-stops⁴, the most effective VR in NIKKOR history, even at super-telephoto distance. Synchro VR, when paired with the Z 9 in-camera VR, effectively compensates up to 6.0-stops. Be it a bird’s colourful plume, or the facial expression of a fox, dive into the exquisite details of your subject with clarity.
Activate SPORT mode
Keep your subjects always in sight with the SPORT mode to effortlessly track fast and erratic movements. The SPORT mode provides a stable image in viewfinder during panning and continuous burst high-speed when you are shooting handheld.
Showcase the finer details
Be impressed by the excellent optical performance of S-Line. Produce high-resolution images from edge to edge with the Super ED and ED glass elements that suppress axial chromatic aberration and with the SR lens element to effectively control short wavelength light with greater precision.
Crystal clear details
See in clarity with a layer of Nano Crystal Coat applied to the front lens element that greatly reduces flare and ghost effect for superb anti-reflective performance.
Well-balanced for handheld shooting.
Hold your lens with greater comfort. Its anti-slip rubber, knurling and small indents on the lens body provide a stable grip when you are shooting handheld. Pan, track, and shoot in comfort with the lens’s centre of gravity positioned close to your body.
Accentuate your subject
Let your subject take centre-stage against a dreamy backdrop. Easily create large and round bokeh with the NIKKOR Z 400mm f/4.5 VR S’s constant maximum aperture of f/4.5, allowing your subject to stand out.
Quiet and accurate AF
Go stealth when photographing wildlife or video recording at a wedding. Unique to Z mount lenses, a stepping motor (STM) that operates quietly and focuses fast is utilised, so you can keep photographing or recording while in a sound-sensitive environment.
Keep shooting, rain or shine
Don’t let the weather hold you back in capturing your favourite subject. The lens is dust- and- drip resistant5, with sealing on various parts of the lens barrel and a rubber gasket on the lens mount, effectively repelling dust and liquid. Remove dirt easily with the fluorine coating applied on the frontmost surface of the lens.
Your preferred setting with one press
The MEMORY SET button conveniently loads your favourite focus position with just one press of a button. Save your frequently used focus position and recall it whenever opportunity calls.